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Modest is an e-mail program specifically targetting modest hardware, such as low-end PCs and internet tablets like the Nokia 770, N800 and N810 internet tablets.
GPE Phone Edition is meant to develop a GTK / GPE based a software stack for mobile Linux devices, especially mobile phones. GPE-Phone's E-mail client is based on the tinymail framework.
TMut is a minimalistic E-mail client. It's the ideal starting point if you want to create a small E-mail client for limited devices. Devices like mobile phones.
There has been talks with the One Laptop Per Child developers about getting tinymail to run on their laptop. As a result, Python bindings have been made. There's also out-of-the-box support for the OLPC environment. The team isn't yet using tinymail, however. Nor has such a decision been made.